Bruce holding Lake Erie Watersnakes
A happy Bruce holding several Lake Erie Watersnakes captured during an annual census. Even snakes have friends.

Dr. Kingsbury and his students have published extensively on issues relating to the ecology and management of a variety of species, and also on the habitats in which they live. The theses of his students also cover a wide variety of topics in their own right – see those here and scroll down to the list of recent graduates and their thesis titles.

Peer-Reviewed Articles

Ratsch, R., B. A. Kingsbury, M. A. Jordan. 2020. Exploration of environmental DNA (eDNA) to detect Kirtland’s Snake (Clonophis kirtlandii). Animals 10:1057.

Tetzlaff, S. J., C. J. Robinson, B. A. Kingsbury, J. H., Sperry; and B. DeGregorio. 2020. Predators May Lose Interest in Turtle Acclimation Pens: Implications for Translocations Using Soft Release. Chelonian Conservation and Biology 19: 141-144.

Tetzlaff, S.J., J.H. Sperry; B. A. Kingsbury and B. DeGregorio. 2019. Captive-rearing duration may be more important than environmental enrichment for enhancing turtle head-starting success. Global Ecology and Conservation 20: https://doi.org/10.1016/j.gecco.2019.e00797.

Degregorio, B.A., M. Ravesi, J. H. Sperry, S. J. Tetzlaff, J. Josimovich, M. Matthews and B. A. Kingsbury. 2018. Daily and seasonal activity patterns of the Massasauga (Sistrurus catenatus): an automated radio-telemetry study. Herpetological Conservation and Biology 13: 10-16.

Josimovich, J., M. Matthews, B. Kingsbury, M. Ravesi. 2018. Sistrurus catenatus (Eastern Massasauga). Defensive behavior. Herpetological Review 49: 355.

Tetzlaff, S. J., M. J. Ravesi, M. C. Allender, E. T. Carter, B. A. DeGregorio, B. A. Kingsbury and J. M. Josimovich. 2017. Snake fungal disease affects behavior of free-ranging Massasauga rattlesnakes. Herpetological Conservation and Biology 12: 624-634.

Tetzlaff, S. J., E. T. Carter, M. J. Ravesi, B. A. DeGregorio and B. A. Kingsbury. 2017. To forage, mate, or thermoregulate: influence of resource manipulation on male rattlesnake behavior. Ecology and Evolution 7: 6606–6613.

Hileman, E. T.  et al. 2017. Climatic and geographic predictors of life history variation in Eastern Massasauga (Sistrurus catenatus): a range-wide synthesis. PLOS ONE https://doi.org/10.1371/journal.pone.0172011.

Matthews, M., J. Josimovich, B. Kingsbury and B. Degregorio. 2017. Sistrurus catenatus (Massasauga). Habitat Use. Herpetological Review 48: 219.

Carter, E.C., M. J. Ravesi, B. C. Eads, B. A. Kingsbury. 2017. Invasive plant management creates ecological traps for snakes. Biological Invasions 19: 443-453.

Ruch, D. G., et al. (multiple authors). 2016. Results of the 2014 Eagle Marsh Biodiversity Survey, Allen County, Indiana. Proceedings of the Indiana Academy of Science 125: 40-49.

Ravesi, M. J., Tetzlaff, S. J., M. C. B. A. Kingsbury. 2016. Detection of Snake Fungal Disease from a Lampropeltis triangulum (Eastern Milksnake) in Northern Michigan. Northeastern Naturalist 23: N18-21.

Attum, O., J. Lowry, B. Kingsbury and E. Carter. 2016. The Distribution of Amur Honeysuckle and Box Turtle Habitat Use in an Urban Forest. Urban Naturalist 12: 1-9.

Ravesi, M., Tetzlaff, S, B. Kingsbury. 2016. Sistrurus catenatus (Massasauga) Diet. Herpetological Review 47: 153-54.

Tetzlaff, S, Ravesi, M., B. Kingsbury. 2016. Sistrurus catenatus (Massasauga) Activity Range Lengths. Herpetological Review 47: 154-5.

Ravesi, M., Forzley, M., Tetzlaff, S, B. Kingsbury. 2015. Sistrurus catenatus (Massasauga) Diet. Herpetological Review 46: 453-454.

Ravesi, M., Tetzlaff, S. J., B. Kingsbury. 2015. Sistrurus catenatus (Massasauga) Diet. Herpetological Review 46: 454.

Carter, E.C., B. C. Eads, M. J. Ravesi, B. A. Kingsbury. 2015. Exotic invasive plants alter thermal regimes: implications for management using a case study of a native ectotherm. Functional Ecology 29: 683-693.

Roe, J. H., M. R. Frank, B. A. Kingsbury. 2015. Altering captive-rearing practices improves success of success of snake reintroductions. Herpetological Conservation and Biology. 10: 711-722.

Tetzlaff, S., Michael Ravesi, Jenna Parker, Michael Forzley, Bruce Kingsbury. 2015. Feeding and breeding: a northern population of Massasauga Rattlesnakes, Sistrurus catenatus (Rafinesque 1818), continues to hunt during the mating season. Herpetology Notes 8: 277-280.

Tetzlaff, S. J., M. Allender, M. Ravesi, J. Smith, B. Kingsbury. 2015. First report of snake fungal disease from Michigan, USA involving Massasaugas, Sistrurus catenatus (Rafinesque 1818). Herpetology Notes 8, 31-33.

Tetzlaff, S.J., Ravesi, M.J., Kingsbury, B.A. 2014. Sistrurus catenatus catenatus (Eastern Massasauga) Diet. Herpetological Review 45(4): 712-713.

Carter, E. T., O. Attum, B. C. Eads, A. S. Hoffman and B. A. Kingsbury. 2014. Reducing the potential for human-snake encounters in a recreational park. Human-Wildlife Interactions 8(2): 158-167.

Roe. J. H, O. Attum and B. A. Kingsbury. 2013. Vital rates and population demographics in declining and stable watersnake populations. Herpetological Conservation and Biology. 8: 591-601.

DeGregorio, B. A. B. Putman and B. A. Kingsbury. 2011. Which habitat selection method is most applicable to Snakes? Case studies of the Eastern Massasauga (Sistrurus catenatus) and Eastern Fox Snake (Pantherophis gloydi) Herpetological Conservation and Biology 6: 372-382.

DeGregorio, B. A. J. V. Manning, N. Bieser and B. A. Kingsbury. 2011. The spatial ecology of the Eastern Massasauga (Sistrurus c. catenatus) in northern Michigan. Herpetologica 67: 71-79.

Roe. J. H, O. Attum and B. A. Kingsbury. 2013. Vital rates and population demographics in declining and stable watersnake populations. Herpetological Conservation and Biology. 8: 591-601.

Jones, P. et al. 2012. Range-wide Analysis of Eastern Massasauga Survival. Journal of Wildlife Management 76: 1576-1586. (Kingsbury and six of his students were among the 32 authors).

DeGregorio, B. A., B. Putman and B. A. Kingsbury. 2011. A Comparison of Macrohabitat Selection Metrics: Which is More Applicable to Snakes? Herpetological Conservation and Biology 6: 372-382.

DeGregorio, B. A. J. V. Manning, N. Bieser and B. A. Kingsbury. 2011. The spatial ecology of the Eastern Massasauga (Sistrurus c. catenatus) in northern Michigan. Herpetologica 67: 71-79.

Roe, J.H., M.R. Frank, S. C. Gibson, O. Attum, and B.A. Kingsbury. 2010. No place like home: an experimental comparison of reintroduction strategies using snakes. Journal of Applied Ecology 47: 1253-1261.

Attum, O., W. Farag, S. M. Baha El Din, and B. Kingsbury. 2010. Retention rate of hard-released translocated Egyptian tortoises Testudo kleinmanni. Endangered Species Research 12:11–15

Attum, O.,  Y. M. Lee, and B. A. Kingsbury. 2009. The Status of the Northern Population of the Copper-bellied Watersnake, Nerodia erythrogaster neglecta. Northeastern Naturalist 16: 317-320.

Marshall, J. C., Jr., B. A. Kingsbury, D. J. Minchella. 2009. Microsatellite Variation, population Structure, and bottlenecks in the threatened copperbelly watersnake. Conservation Genetics 10: 465-476.

Attum, O., B. Rabea, S. Osman, S. Habinan, S. Baha El Din, B. Kingsbury. 2008. Conserving and tracking tortoises: A local community visual tracking or radio-tracking approach? Journal of Arid Environments 72: 671-676.

Attum, O., Y. M. Lee, J. H. Roe and B. A. Kingsbury. 2008. Wetland complexes and upland–wetland linkages: landscape effects on the distribution of rare and common wetland reptiles. Journal of Zoology 275: 245-251.

Cooper, W. E., O. Attum, and B. Kingsbury. 2008. Escape behaviors and flight initiation distance in the common water snake Nerodia sipedon. Journal of Herpetology 42: 493-500.

Attum, O., M. Esawy, W. Farag, A. Gad, S. Baha El Din, B. Kingsbury. 2007. Returning them back to the wild: Movement patterns of repatriated Egyptian tortoises. Zoology in the Middle East 41: 35-40.

Attum, O., Y. M. Lee, J. H. Roe and B. A. Kingsbury. 2007. Upland–wetland linkages: relationship of upland and wetland characteristics with watersnake abundance. Journal of Zoology 271: 134–139.

Attum, O., S. Baha El Din, S. Carranza, R. Earley, E.N. Arnold, and B. Kingsbury.  2007. An evaluation of the taxonomic validity of Testudo werneri. Amphibia-Reptilia 28: 393-401.

Roe, J. H., J. K. Gibson, and B. A. Kingsbury. 2006. Beyond the wetland border: estimating the impact of roads for two species of watersnakes. Biological Conservation 130: 161-168.

Marshall, J. C., Jr., J. M. Manning, and B. A. Kingsbury. 2006. Movement and macrohabitat selection of the Eastern Massasauga in a fen habitat. Herpetologica 62: 141-150.

Roe, J. H., B. A. Kingsbury, and M. A. Myers. 2005. Nerodia erythrogaster neglecta (Copper-bellied Watersnake) foraging behavior. Herpetological Review 36: 70.

Roe, J. H., B. A. Kingsbury, N. R. Herbert. 2004. Comparative water snake ecology: conservation of mobile animals that use temporally dynamic resources. Biological Conservation 118: 79-89.

Roe, J. H., B. A. Kingsbury, N. R. Herbert. 2003. Wetland and upland use patterns in the semi-aquatic snakes: implications for wetland conservation. Wetlands 23(4): 1003-1014.

Laurent, E. J., and B. A. Kingsbury. 2003. Habitat separation between three species of water snake in northwestern Kentucky, including the endangered copperbelly water snake. Journal of Herpetology 37 (2): 229-235.

Kingsbury, B. A. and C. J. Coppola. 2000. Hibernacula of the copperbelly water snake (Nerodia erythrogaster neglecta) in southern Indiana and Kentucky. Journal of Herpetology 34: 294-298.

Petzing, J. E., M. J. Dreslik, C. A. Phillips, C. D. Smith, A. R. Kunhs , D. B. Shepard, J. G. Palis, E. O.  Moll, D. J. Olson, T. G. Anton, D. Mauger, and B. Kingsbury. 2000. New amphibian and reptile county records for Illinois.  Herpetological Review. 31(3)189-194.

Kingsbury, B. A. 1999. An experimental design for examining thermoregulatory set points in ectothermic animals. The American Biology Teacher 61: 448-452.

Kingsbury, B. A. 1996. Status of the eastern massasauga, Sistrurus catenatus catenatus, in Indiana, with management recommendations for recovery. Proceedings of the Indiana Academy of Sciences 105: 195-205.

Kingsbury, B. A. 1995. Field metabolic rates of a eurythermic lizard. Herpetologica 51: 155-159.

Kingsbury, B. A. 1994. Thermal constraints and eurythermy in the lizard Elgaria multicarinata. Herpetologica 50: 266-273.

Kingsbury, B. A. 1993. Thermoregulatory set points of the eurythermic lizard Elgaria multicarinata. Journal of Herpetology 27: 241-247.

Books and Book Chapters

Kingsbury, B. A.  In progress. A Field Guide to the Reptiles and Amphibians of Indiana. Contracted with Indiana University Press, Bloomington, Indiana.

Kingsbury, B. A. Accepted with revision. Copper-bellied Watersnake. In: Davis JD, Lipps GJ, Matson TO, Pfingsten RA, Wynn DE (editors).  Amphibians and Reptiles of Ohio.  Ohio Biological Survey, Columbus, Ohio, USA.  Invited book chapter.

Kingsbury, B. A. and D. Keyler. In press. Indiana. In: The Timber Rattlesnake Conservation Action Plan. Partners in Amphibian and Reptile Conservation.

Kingsbury, B. A., and N. J. Robinson. 2016. Movement patterns and telemetry. In: Reptile Ecology and Conservation: A Handbook of Techniques, C. K. Dodd, ed. Oxford University Press, Oxford, UK.

Whitaker, Jr., J. O, and C. J. Amlaner, eds. 2012. Habitats and Ecological Communities of Indiana: Pre-settlement to Present. J.O. Whitaker, Jr., and C. J. Amlaner, eds.  Indiana University Press, Bloomington, Indiana. 512 pp. I was the herpetologist on the team and wrote content throughout the book.

Kingsbury, B. A. and J. Gibson, eds. 2012. “Habitat Management Guidelines for Amphibians and Reptiles of the Midwest”, 2nd ed. Partners in Amphibian and Reptile Conservation. 140 pp.

Kingsbury, B. A. and O. Attum. 2009. Conservation Strategies: Captive breeding, translocation, and repatriation. In: Snakes: Ecology and Conservation, S. Mullin and R. Seigel, eds. Cornell University Press.  365 pp.

Jenkins, C., C. R., Peterson and B. A. Kingsbury. 2009. Modeling Snake Distribution and Habitat. In: Snakes: Ecology and Conservation, S. Mullin and R. Seigel, eds. Cornell University Press. 365 pp.

Gibson, S. E., Z. J. Walker, and B. A. Kingsbury. 2008. Microhabitat preferences of the Timber Rattlesnake (Crotalus horridus) in the Hardwood Forests of Southern Indiana. In: Biology of the Rattlesnakes, K. R. Beaman, S. P. Bush, M. D. Cardwell, and W. K. Hayes, eds., pp. 275-286.

Williams, R. N., B. MacGowan, B. A. Kingsbury, and Z. J. Walker. 2006. “Salamanders of Indiana.” Purdue University Extension and the Indiana Department of Natural Resources.

MacGowan, B., B. A. Kingsbury, and R. N. Williams. 2005. “Turtles of Indiana” Purdue University Extension and the Indiana Department of Natural Resources. 63 pp.

Kingsbury, B. A., J. Gibson, and contributors. 2002. “Habitat Management Guidelines for Amphibians and Reptiles of the Midwest.” 57 pp.

MacGowan, B. and B. A. Kingsbury. 2001. “Snakes of Indiana.” Published by Purdue University Extension and the Indiana Department of Natural Resources. 51 pp.

Other Publications of Interest

Kingsbury, B. A. and C. A. Tansy. 2008. Copperbelly Water Snake (Nerodia erythrogaster neglecta) Recovery Plan. US Fish and Wildlife Service. 58 pp. plus appendices.

Tansy, C. and B. Kingsbury. 2005. Copperbelly Water Snake (Northern Population) Recovery Outline. U.S. Fish and Wildlife Service. 17 pp.

Gibson, J. K. and B. A. Kingsbury. 2004. Conservation Assessment for Kirtland’s Snake (Clonophis kirtlandii). USDA Forest Service. 29 pp.

MacGowan, B. and B. A. Kingsbury. 2004. “Snakes of the Midwest.” An interactive CD published by Purdue University Extension and the Center for Reptile and Amphibian Conservation and Management.

Kingsbury, B. A. 2002. R9 Conservation Approach for the Eastern Massasauga (Sistrurus c. catenatus) on the Huron-Manistee National Forests. US Forest Service.

Kingsbury, B. A. and R. Grimes. 2001. “A Description and Assessment of the Copperbelly Water Snake Conservation Agreement.” Transactions of the Sixty-sixth North American Wildlife and Natural Resources Conference: pp. 538-554.

Johnson, G., R. King, B. Kingsbury, C. Parent, R. Seigel, and J Szymanski. 2000. “The Eastern Massasauga: A Handbook for Land Managers.” US Fish and Wildlife Service.

Kingsbury, B. A. 1999. Promoting Massasauga Conservation in Public Wildlife Areas. Proceedings of the Second International Symposium and Workshop on the Conservation of the Eastern Massasauga Rattlesnake, Sistrurus c. catenatus. Toronto Metropolitan Zoo.